Vernonia glaberrima

Vernonia glaberrima

Gymnanthemum glaberrimum

Common Names: 
Local Names: Shiwakaar, Shiwaakar-jan-ga gari (Hausa, Nigeria)
Species ID: NMP-285 |
wfo-0000134063

Scientific Classification

Kingdom: Plantae

Clade: Tracheophytes

Clade: Angiosperms

Clade: Eudicots

Clade: Asterids

Order: Asterales

Family: Asteraceae

Genus: Gymnanthemum

Species: Gymnanthemum glaberrimum (Welw. ex O.Hoffm.) H. Rob.

Synonyms: Vernonia glaberrima Welw. ex O.Hoffm.; Vernonia hensii Klatt (treated as a synonym in some African floras).

Morphological Description

Gymnanthemum glaberrimum is a much-branched, perennial subshrub or small shrub with a woody base, usually 0.5–1 m tall but occasionally reaching about 2 m in favourable sites. It often forms small, open tufts in grassland and savanna, becoming conspicuous when in full flower.

·       Stems: Slender, erect to ascending, terete, glabrous to sparsely pubescent. Plants are characteristically almost leafless at peak flowering, with older leaves shed and new ones emerging shortly after, giving a delicate, twiggy aspect to flowering stands (Isawumi, 1995; Hyde et al., 2025).

·       Leaves: Elliptic to oblanceolate, about 3–10 cm long and 1–3 cm wide, with subentire to coarsely dentate-serrate margins. Apex acute to acuminate, base cuneate; texture coriaceous and often somewhat shiny; surfaces mostly glabrous and dark green. In some floristic treatments, leaves are described as appearing shortly after flowering, suggesting adaptation to seasonal climates ( Hyde et al., 2025).

·       Inflorescences: Numerous small capitula, usually 5–8 mm in diameter, borne in open panicles or pseudumbels on short lateral branches. Florets are white, tubular, sweetly scented and five-lobed; involucral bracts are several-seriate and narrowly lanceolate. Flowering is commonly observed in the dry season or at the end of the rains in many parts of its range.

·       Fruit: Cylindrical achenes, about 2–3 mm long, 10-ribbed, crowned by a pappus of white bristles. The light achenes are wind-dispersed and can rapidly colonise disturbed patches (Fern, 2024; Hyde et al., 2025).

·       Microscopic characters: Pharmacognostic studies on the leaf report amphistomatic blades with anomocytic stomata, stomatal indices around 3–4% (adaxial) and 11% (abaxial), and the presence of oil glands, starch grains, tannins, cellulose, gum, calcium oxalate crystals, fibres and vessel elements.

Distribution and Habitat

Gymnanthemum glaberrimum is African in distribution and corresponds to Vernonia glaberrima in older floras.

·       Geographic range: West Africa (from Guinea through Togo and Nigeria to Cameroon), Central Africa (Democratic Republic of Congo), and Eastern to Southern Africa, including Tanzania, Malawi, Zambia and Zimbabwe (Kew Science, n.d.; Hyde et al., 2025).

·       Altitude: Typically recorded between 600 and 1,600 m above sea level in Flora of Tropical East Africa and Flora Zambesiaca accounts (Kew Science, n.d.; Flora of Zambia, 2025).

·       Habitats: Open hillside grasslands, wooded grassland and Brachystegia-dominated miombo woodland, forest margins, fallow fields and other disturbed sites. Often locally frequent in well-drained, sandy or loamy soils with moderate to high rainfall (c. 600–1,800 mm).

·       Regional notes: In Zimbabwe and Malawi it is described as a frequent native shrub of grassland and woodland edges, sometimes forming scattered clumps in open savanna (Hyde et al., 2025). In Nigeria it occurs mainly in northern savanna zones and forest–savanna transition belts, often on fallow land and roadsides.

·       Ecological role: A light-demanding pioneer species that colonises disturbed ground, providing nectar and pollen for insects (bees, butterflies and other pollinators). The root system helps stabilise soil on slopes and contributes to herbaceous diversity in savannas and open woodland.

Ethnopharmacology

Under its synonym Vernonia glaberrima, this species is recognised in West and Central African ethnomedicine. Ethnobotanical surveys and pharmacological introductions consistently report its use for inflammatory and infectious conditions, malaria, diabetes and skin-related disorders, particularly skin cancers and chronic lesions (Alhassan et al., 2018; Abdullahi et al., 2015a; Gangas et al., 2021; Onugwu et al., 2024).

·       Malaria and febrile illness: Decoctions of the leaves are taken orally in northern Nigeria and neighbouring regions to treat malaria and fever, often in combination with other antipyretic herbs. This use is explicitly mentioned in several pharmacological papers and reviews of the species (Abdullahi et al., 2015b; de Villiers et al., 2010; Gangas et al., 2021).

·       Skin cancer and chronic skin conditions: Leaf preparations, especially methanolic or aqueous extracts and poultices, are used traditionally in West Africa for the management of skin cancers, chronic skin ulcers and other skin-related disorders (Alhassan et al., 2018; Abubakar et al., 2022).

·       Pain and inflammation: The plant is widely cited in local practice as a remedy for body pains, headaches, arthritis and general inflammatory complaints. Leaves may be taken as decoctions or used topically as poultices (Abdullahi et al., 2016; Yusuf et al., 2020).

·       Infectious and gastrointestinal diseases: Traditional healers use leaf extracts for diarrhoea, gastrointestinal discomfort and a range of microbial infections, including wound and skin infections. These uses align with observed antibacterial activity in vitro (Abdullahi et al., 2015c; Gangas et al., 2021).

·       Diabetes and metabolic disorders: In northern Nigeria, leaf infusions or decoctions are administered to people with “sugar disease” (diabetes mellitus), reflecting its hypoglycaemic potential demonstrated in experimental models (Abdullahi et al., 2015a).

·       Mosquito-borne disease control: Recent ethnopharmacological-inspired work has explored the plant as a larvicidal and mosquito-repellent agent against Aedes mosquitoes, supporting its emerging role in community-based vector control strategies (Onugwu et al., 2024).

A growing body of in vitro and in vivo work substantiates many of the traditional uses of Gymnanthemum glaberrimum, especially under its synonym Vernonia glaberrima. Key activities are summarised below.

Antimicrobial and Antibacterial Activity

·       Crude methanol leaf extract showed broad-spectrum antimicrobial activity against clinically important pathogens, including methicillin-resistant Staphylococcus aureus, vancomycin-resistant enterococci, Listeria monocytogenesHelicobacter pylori, various Gram-negative bacteria and pathogenic yeasts, using agar diffusion and broth dilution methods (Abdullahi et al., 2015c; Abdullahi et al., 2015d).

·       Hexane extracts of leaves and stems, alone and in combination with standard antibiotics, displayed notable antibacterial activity and synergistic effects against multidrug-resistant bacteria. These findings support the traditional use of the plant for infectious and skin conditions (Gangas et al., 2021).

Antimalarial Activity

·       Methanolic leaf extract produced significant, dose-dependent suppression of parasitaemia in mice infected with chloroquine-sensitive Plasmodium berghei (NK65) in a curative model, confirming antiplasmodial activity that aligns closely with folk use for malaria (Abdullahi et al., 2015b; de Villiers et al., 2010).

Antidiabetic and Metabolic Effects

·       In alloxan-induced diabetic rats, methanolic leaf extract produced a significant, time-dependent reduction in fasting blood glucose, improvement in glucose tolerance and a moderate hypoglycaemic effect overall. The study concluded that the extract may be useful as an adjunct in diabetes management (Abdullahi et al., 2015a).

·       Subsequent reviews on Vernonia species have highlighted V. glaberrima as one of several antidiabetic Vernonieae taxa, often linked to the presence of triterpenes and flavonoids (Yusuf et al., 2024).

Analgesic and Anti-inflammatory Activity

·       The n-hexane-soluble fraction of methanolic leaf extract significantly reduced acetic acid–induced writhing in mice and carrageenan-induced paw oedema in rats, demonstrating both peripheral analgesic and anti-inflammatory effects and experimentally validating traditional use for pain and swelling (Abdullahi et al., 2016).

·       Lupeol isolated from the n-hexane fraction exhibited strong analgesic activity in acetic acid–induced writhing tests (over 80% inhibition at doses of 12.5–50 mg/kg) and significantly attenuated both early and late phases of formalin-induced pain, as well as formalin-induced paw inflammation. The median LD50 for lupeol was greater than 5,000 mg/kg, indicating low acute toxicity (Yusuf et al., 2020).

·       A hydroethanolic leaf extract displayed anti-inflammatory and immunomodulatory effects, reducing protein denaturation, stabilising lysosomal membranes and modulating cyclophosphamide-induced immunosuppression in experimental models (Ibrahim et al., 2025).

Anticancer and Cytotoxic Activity

·       Methanolic and chloroform leaf extracts and their purified fractions showed significant cytotoxicity against multiple human cancer cell lines, including breast (MCF7), cervical (HeLa), liver (HepG2), colon and melanoma cells. Isolated compounds such as lupeol and 5-methylcoumarin-4-β-glucoside contributed strongly to these effects (Alhassan et al., 2018; Abubakar et al., 2022).

·       Molecular docking and in vitro assays implicate targets such as cyclin-dependent kinase 2 (CDK2) and carbonic anhydrase IX (CAIX) in the anticancer effects, providing mechanistic support for the traditional use of the plant against skin cancers (Alhassan et al., 2018).

Mosquito Larvicidal and Repellent Activity

·       Methanolic leaf extract and fractions exhibited potent larvicidal activity against Aedes aegypti larvae, with aqueous methanol fractions showing LC50 values as low as approximately 2.8 ppm and sub-fractions achieving up to 75% larval mortality. Phytochemical analyses associated these effects with flavonoids, terpenoids, saponins and phenolics (Onugwu et al., 2024).

·       Leaf extract fractions also demonstrated strong mosquito-repellent activity against adult Aedes aegypti, with percentage repellency approaching 99% at the highest tested concentrations, suggesting promising application as a plant-based mosquito repellent for malaria and arboviral disease control.

Antivenom-related Potential

·       Computational docking and experimental validation studies on selected leaf phytochemicals from V. glaberrima showed inhibitory potential against key snake venom toxins, including phospholipase A2 and metalloproteinases, and reduced venom-induced cellular damage in vitro. These findings point to a future role for the plant as an adjunct in antivenom therapy (Yusuf et al., 2024).

⚠ Safety Considerations: Experimental studies suggest that leaf extracts of Gymnanthemum glaberrimum are generally well tolerated at doses showing antimalarial, antidiabetic, analgesic and anti-inflammatory effects. Nonetheless, high or prolonged dosing can alter liver and kidney parameters in animals, and long-term, high-dose human use has not been rigorously evaluated. Use should remain within traditional dose ranges and under guidance of trained practitioners, especially in individuals with underlying hepatic or renal disease, during pregnancy or in children.

Additional Notes

·       Taxonomic reclassification: Modern systematic work places Vernonia glaberrima within Gymnanthemum, leading to the currently accepted name Gymnanthemum glaberrimum. This is reflected in Kew’s Plants of the World Online and the World Flora Online (Kew Science, n.d.).

·       Conservation: The species is generally common in suitable habitats across several countries and is not currently listed as threatened in regional floras. However, local populations may be impacted by agricultural expansion, overgrazing and frequent burning.

·       Research gaps: Although considerable progress has been made in elucidating its phytochemistry and preclinical pharmacology, there is still limited information on pharmacokinetics, herb–drug interactions and well-designed clinical trials. Standardised extracts and clear dosing guidelines would support future clinical development.

References

  • Abdullahi, M. I., Uba, A., Yaro, A. H., Maxwell, O., Kabir, S., Alhasan, A. M., Umar, A., Bello, S. S., & Nasir, I. (2015a). Phytochemical screening, acute toxicity study and evaluation of antidiabetic properties of the methanolic leaf extract of Vernonia glaberrima (Asteraceae). Journal of Pharmaceutical, Chemical and Biological Sciences3(2), 169–177.
  • Abdullahi, M. I., Uba, A., Yusuf, A. J., Nasir, I., Muntaka, A., Alhasan, A. M., Alebiosu, C. O., Yahaya, M., Yunusa, S. D., & Sadiq, M. (2016). Phytochemical analysis, analgesic and anti-inflammatory studies on the n-hexane soluble fraction of Vernonia glaberrima. International Journal of Science for Global Sustainability2(4), 16–23.
  • Abdullahi, M. I., Uba, A., Alebiosu, C. O., Alhasan, A. M., Mode, S., Umar, A., Nasir, I., Bello, S. S., Ibrahim, Z. Y. Y., Yunusa, A., & Yusuf, A. J. (2015b). Evaluation of the antimalarial effect of the methanolic leaf extract of Vernonia glaberrima (Asteraceae) against Plasmodium berghei infected mice. IOSR Journal of Pharmacy and Biological Sciences10(1), 6–9.
  • Abdullahi, M. I., Uba, A., Alebiosu, C. O., Alhasan, A. M., Mode, S., Umar, A., Nasir, I., Bello, S. S., Ibrahim, Z. Y. Y., Yunusa, A., & Yusuf, A. J. (2015c). Antimicrobial properties of the methanol leaf extract of Vernonia glaberrima Welw. ex O. Hoffm (Asteraceae). African Journal of Pharmacy and Pharmacology9(37), 914–918.
  • Abu, D. E., Tim, J. N., Imo-Obasi, P., & Ayoka, T. O. (2022). In-vivo sub-chronic toxicological evaluation of extract of Vernonia glaberrima leaves in experimental rats. Notulae Scientia Biologicae14(2), 11181.
  • Abubakar, B., Alhassan, A. M., Malami, I., & Imam, M. U. (2022). Evaluation of acute and sub-acute toxicity profile of 5-methylcoumarin-4β-glucoside in mice, a chemotherapeutic candidate isolated from Vernonia glaberrima leaves. Toxicology Reports9, 215–225.
  • Alhassan, A. M., Ahmed, Q. U., Latip, J., Shah, S. A. A., Sarian, M. N., Wahab, R. A., Taher, M., Abdullahi, M. I., & Khatib, A. (2018). Phytoconstituents from Vernonia glaberrima Welw. ex O. Hoffm. leaves and their cytotoxic activities on a panel of human cancer cell lines. South African Journal of Botany116, 16–24.
  • de Villiers, B. J., Van Vuuren, S. F., Van Zyl, R. L., & Van Heerden, F. R. (2010). Antimicrobial and antimalarial activity of Cussonia species and related triterpenes. Journal of Ethnopharmacology129(2), 189–196.
  • Flora of Zambia. (2025). Vernonia glaberrima (syn. Gymnanthemum glaberrimum) species information. In Flora of Zambia. https://www.zambiaflora.com/speciesdata/species.php?species_id=158290
  • Gangas, P., Dangoggo, S. M., Yauri, U. A., & co-authors. (2021). GC–MS analysis and antibacterial effects of Vernonia glaberrima n-hexane extracts alone and in combination with standard antibiotics. Journal of the Chemical Society of Nigeria46(3), 303–314.
  • Hyde, M. A., Wursten, B. T., Ballings, P., & Coates Palgrave, M. (2025). Vernonia glaberrima Welw. ex O.Hoffm. In Flora of Zimbabwe. https://www.zimbabweflora.co.zw/speciesdata/species.php?species_id=158290
  • Ibrahim, S. B., & co-authors. (2025). Immunomodulatory and anti-inflammatory properties of the hydroethanolic extract of Vernonia glaberrima. African Journal of Pharmacy and Pharmacology19(3), 145–156.
  • Kew Science. (n.d.). Gymnanthemum glaberrimum (Welw. ex O.Hoffm.) H. Rob. Plants of the World Online. Royal Botanic Gardens, Kew. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:258418-1
  • Onugwu, S. O., Onugwu, A. L., & Ezugwu, C. O. (2024). Phytochemical and larvicidal evaluation of Vernonia glaberrima Welw. ex O. Hoffm. (Asteraceae) leaves on Aedes mosquito larvae. Trends in Natural Products Research5(2), 81–90.
  • Yusuf, A. J., Bugaje, A. I., Sadiq, M., Salihu, M., Adamu, H. W., & Abdulrahman, M. (2024). Exploring the inhibitory potential of phytochemicals from Vernonia glaberrima leaves against snake venom toxins through computational simulation and experimental validation. Toxicon247, 107838.
  • Yusuf, A. J., Jega, A. Y., & co-authors. (2020). Effect of lupeol from Vernonia glaberrima (Asteraceae) on pain and inflammation. Journal of Medicinal Plants for Economic Development4(1), 84.

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Compounds of Vernonia glaberrima
References

Alhassan, A. M., Ahmed, Q. U., Latip, J., Shah, S. A. A., Sarian, M. N., Wahab, R. A., ... & Khatib, A. (2018). Phytoconstituents from Vernonia glaberrima Welw. Ex O. Hoffm. leaves and their cytotoxic activities on a panel of human cancer cell lines. South African journal of botany, 116, 16-24.